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bh, ch, fh, gh, mh, ph, sh were pronounced as today. bh, ch, gh, mh, ph were pronounced like b, c, g, m, p but without an oral stop of air flow, i.e. as corresponding fricatives. sh as /h/, fh silent because s and f are already fricatives. They lost frication. mh was a nasal /ṽ/, bh a simple /v/. (broad /w/ is probably a later innovation)
dh, th were pronounced as d, t without oral stop of air flow, that means as dental fricatives, like English th (dh voiced /ð/, th voiceless /θ/). There were probably different broad and slender versions, too.
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